I was reading O’Donnell, Liza et al. “Spermiation: The Process of Sperm Release.” and got stuck with a question.
From the paper,'' Spermiation is the process by which mature spermatids are released from Sertoli cells into the seminiferous tubule lumen prior to their passage to the epididymis.''
When did the spermatids got attached/associated to Sertoli cells that during spermiation they were released from the sertoli cells?